Theory of multicellularity Emergence of life cycles From a solitary cell to a colony Evolution of cell differentiation From a colony of identical cells to a collective with diverse members Eco-evolutionary dynamics Evolution of migration They move Evolution of interactions From antagonism to cooperation and beyond Synergies with experiments (empirical collaborations) Empirical evolution of multicellularity The theory brought to the lab Oncology ecDNA dynamics Morphology Skulls of legless lizards